The lightning mole is a medium-sized subterranean animal featured in The Precursor Legacy. They were featured in the mission "Herd the moles into their hole", during which Jak had to chase them back into the holes they were scared out of by a band of robbers in the Precursor Basin. They were discovered and researched by the Geologist, who rewarded Jak and Daxter a power cell for herding the moles.
Lightning moles, despite their name, hardly resemble moles as much as they resemble squirrels. They have orange, velvety fur, and large dark eyes. Their ability to run quite fast comes from their strong hind legs. They are blind, and rely on their sense of hearing to traverse the foreign Precursor Basin platforms.